Stan Wullschleger

First name
Last name
Project Leadership Role(s)
  • Management
  • Special Project Advisor
Project Leadership Title(s)
  • Special Project Advisor


  • Conroy, N. A., et al. “Chemostatic Concentration–Discharge Behaviour Observed In A Headwater Catchment Underlain With Discontinuous Permafrost”. Hydrological Processes, 2022.
  • McCaully, R. E., et al. “High Temporal And Spatial Variability Of Nitrate On An Alaskan Hillslope Dominated By Alder Shrubs”. The Cryosphere, 2022.
  • Arendt, C. A., et al. “Increased Arctic No3− Availability As A Hydrogeomorphic Consequence Of Permafrost Degradation And Landscape Drying”. Nitrogen, 2022, pp. 314 - 332.
  • Zheng, J., et al. “Quantifying Ph Buffering Capacity In Acidic, Organic-Rich Arctic Soils: Measurable Proxies And Implications For Soil Carbon Degradation”. Geoderma, 2022, p. 116003.
  • Curasi, S. R., et al. “Range Shifts In A Foundation Sedge Potentially Induce Large Arctic Ecosystem Carbon Losses And Gains”. Environmental Research Letters, 2022, p. 045024.
  • Bennett, K. E., et al. “Spatial Patterns Of Snow Distribution For Improved Earth System Modelling In The Arctic”. The Cryosphere, 2022.
  • Zhang, L., et al. “Unravelling Biogeochemical Drivers Of Methylmercury Production In An Arctic Fen Soil And A Bog Soil”. Environmental Pollution, 2022, p. 118878.


  • Ely, K. S., et al. “A Reporting Format For Leaf-Level Gas Exchange Data And Metadata”. Ecological Informatics, 2021, p. 101232.
  • Roy_Chowdhury, T., et al. “Temporal, Spatial, And Temperature Controls On Organic Carbon Mineralization And Methanogenesis In Arctic High-Centered Polygon Soilsdata_Sheet_1.Docx”. Frontiers In Microbiology, 2021.
  • Ladd, M. P., et al. “Untargeted Exometabolomics Provides A Powerful Approach To Investigate Biogeochemical Hotspots With Vegetation And Polygon Type In Arctic Tundra Soils”. Soil Systems, 2021, p. 10.


  • Philben, M. J., et al. “Anaerobic Respiration Pathways And Response To Increased Substrate Availability Of Arctic Wetland Soils”. Environmental Science: Processes & Impacts, 2020, pp. 2070 - 2083.
  • Iversen, C. M., et al. “Building A Culture Of Safety And Trust In Team Science”. Eos, 2020.
  • Krassovski, M. B., et al. “Hybrid-Energy Module For Remote Environmental Observations, Instruments, And Communications”. Advances In Polar Science , 2020, pp. 156-166.
  • Philben, M. J., et al. “Influences Of Hillslope Biogeochemistry On Anaerobic Soil Organic Matter Decomposition In A Tundra Watershed”. Journal Of Geophysical Research: Biogeosciences, 2020.
  • Herndon, E. M., et al. “Iron And Iron-Bound Phosphate Accumulate In Surface Soils Of Ice-Wedge Polygons In Arctic Tundra”. Environmental Science: Processes & Impacts, 2020, pp. 1475 - 1490.
  • Lara, M. J., et al. “Local-Scale Arctic Tundra Heterogeneity Affects Regional-Scale Carbon Dynamics”. Nature Communications, 2020.
  • Conroy, N. A., et al. “Timing And Duration Of Hydrological Transitions In Arctic Polygonal Ground From Stable Isotopes”. Hydrological Processes, 2020, pp. 749-764.
  • Conroy, N. A., et al. “Timing And Duration Of Hydrological Transitions In Arctic Polygonal Ground From Stable Isotopes”. Hydrological Processes, 2020, pp. 749 - 764.
  • Wales, N. A., et al. “Understanding The Relative Importance Of Vertical And Horizontal Flow In Ice-Wedge Polygons”. Hydrology And Earth System Sciences, 2020, pp. 1109-1129.


  • Salmon, V. G., et al. “Alder Distribution And Expansion Across A Tundra Hillslope: Implications For Local N Cycling”. Frontiers In Plant Science, 2019.
  • Ladd, M. P., et al. “Evaluation Of An Untargeted Nano-Liquid Chromatography-Mass Spectrometry Approach To Expand Coverage Of Low Molecular Weight Dissolved Organic Matter In Arctic Soil”. Scientific Reports, 2019.
  • Herndon, E. M., et al. “Iron (Oxyhydr)Oxides Serve As Phosphate Traps In Tundra And Boreal Peat Soils”. Journal Of Geophysical Research: Biogeosciences, 2019, pp. 227 - 246.
  • Wang, Y., et al. “Mechanistic Modeling Of Microtopographic Impacts On Carbon Dioxide And Methane Fluxes In An Alaskan Tundra Ecosystem Using The Clm‐Microbe Model”. Journal Of Advances In Modeling Earth Systems, 2019, p. 17.
  • Zheng, J., et al. “Modeling Anaerobic Soil Organic Carbon Decomposition In Arctic Polygon Tundra: Insights Into Soil Geochemical Influences On Carbon Mineralization”. Biogeosciences, 2019, pp. 663 - 680.
  • Yang, Z., et al. “Temperature Sensitivity Of Mineral-Enzyme Interactions On The Hydrolysis Of Cellobiose And Indican By Beta-Glucosidase”. Science Of The Total Environment, 2019, pp. 1194 - 1201.
  • Rogers, A., et al. “Terrestrial Biosphere Models May Overestimate Arctic Carbon Dioxide Assimilation If They Do Not Account For Decreased Quantum Yield And Convexity At Low Temperature”. New Phytologist, 2019, pp. 167 - 179.


  • Jubb, A. M., et al. “Characterization Of Iron Oxide Nanoparticle Films At The Air–Water Interface In Arctic Tundra Waters”. Science Of The Total Environment, 2018, pp. 1460-1468.
  • Young-Robertson, J. M., et al. “Evaporation Dominates Evapotranspiration On Alaska’s Arctic Coastal Plain”. Arctic, Antarctic, And Alpine Research, 2018, p. e1435931.
  • Zheng, J., et al. “Impacts Of Temperature And Soil Characteristics On Methane Production And Oxidation In Arctic Polygonal Tundra”. Biogeosciences Discussions, 2018, pp. 1 - 27.
  • Fisher, J. B., et al. “Missing Pieces To Modeling The Arctic-Boreal Puzzle”. Environmental Research Letters, 2018, p. 020202.
  • Chen, H., et al. “Molecular Insights Into Arctic Soil Organic Matter Degradation Under Warming”. Environmental Science & Technology, 2018, pp. 4555-4564.


  • Raz-Yaseef, N., et al. “Evapotranspiration Across Plant Types And Geomorphological Units In Polygonal Arctic Tundra”. Journal Of Hydrology, 2017, pp. 816-825.
  • Raz-Yaseef, N., et al. “Large Carbon Dioxide And Methane Emissions From Polygonal Tundra During Spring Thaw In Northern Alaska”. Geophysical Research Letters, 2017, pp. 504 - 513.
  • Yang, Z., et al. “Microbial Community And Functional Gene Changes In Arctic Tundra Soils In A Microcosm Warming Experiment”. Frontiers In Microbiology, 2017.
  • Wullschleger, S. D. “Profile: Stan D. Wullschleger”. New Phytologist, 2017, pp. 981 - 983.
  • Rogers, A., et al. “Terrestrial Biosphere Models Underestimate Photosynthetic Capacity And Carbon Dioxide Assimilation In The Arctic”. New Phytologist, 2017, pp. 1090 - 1103.
  • Walker, A. P., et al. “Trait Covariance: The Functional Warp Of Plant Diversity?”. New Phytologist, 2017, pp. 976-980.


  • Ali, A. A., et al. “A Global Scale Mechanistic Model Of Photosynthetic Capacity (Luna V1.0)”. Geoscientific Model Development, 2016, pp. 587 - 606.
  • Throckmorton, H. M., et al. “Active Layer Hydrology In An Arctic Tundra Ecosystem: Quantifying Water Sources And Cycling Using Water Stable Isotopes”. Hydrological Processes, 2016.
  • Yang, Z., et al. “Effects Of Warming On The Degradation And Production Of Low-Molecular-Weight Labile Organic Carbon In An Arctic Tundra Soil”. Soil Biology And Biochemistry, 2016, pp. 202 - 211.
  • Langford, Z. L., et al. “Mapping Arctic Plant Functional Type Distributions In The Barrow Environmental Observatory Using Worldview-2 And Lidar Datasets”. Remote Sensing, 2016, p. 733.
  • Xu, X., et al. “Reviews And Syntheses: Four Decades Of Modeling Methane Cycling In Terrestrial Ecosystems”. Biogeosciences, 2016, pp. 3735 - 3755.
  • Yang, Z., et al. “Warming Increases Methylmercury Production In An Arctic Soil”. Environmental Pollution, 2016, pp. 504 - 509.


  • Xu, X., et al. “A Microbial Functional Group-Based Module For Simulating Methane Production And Consumption: Application To An Incubated Permafrost Soil”. Journal Of Geophysical Research: Biogeosciences, 2015, pp. 1315 - 1333.
  • Wullschleger, S. D., et al. “Genomics In A Changing Arctic: Critical Questions Await The Molecular Ecologist”. Molecular Ecology, 2015, pp. 2301 - 2309.
  • Herndon, E. M., et al. “Geochemical Drivers Of Organic Matter Decomposition In Arctic Tundra Soils”. Biogeochemistry, 2015, pp. 397 - 414.
  • Ali, A. A., et al. “Global-Scale Environmental Control Of Plant Photosynthetic Capacity”. Ecological Applications, 2015, pp. 2349 - 2365.
  • Mann, B. F., et al. “Indexing Permafrost Soil Organic Matter Degradation Using High-Resolution Mass Spectrometry”. Plos One, 2015.
  • Heikoop, J. M., et al. “Isotopic Identification Of Soil And Permafrost Nitrate Sources In An Arctic Tundra Ecosystem”. Journal Of Geophysical Research: Biogeosciences, 2015, pp. 1000 - 1017.
  • Wullschleger, S. D., et al. “Leaf Respiration (Globresp) - Global Trait Database Supports Earth System Models”. New Phytologist, 2015, pp. 483 - 485.
  • Cohen, L. R., et al. “Measuring Diurnal Cycles Of Evapotranspiration In The Arctic With An Automated Chamber System”. Ecohydrology, 2015, pp. 652 - 659.
  • Newman, B. D., et al. “Microtopographic And Depth Controls On Active Layer Chemistry In Arctic Polygonal Ground”. Geophysical Research Letters, 2015, pp. 1808 - 1817.
  • Throckmorton, H. M., et al. “Pathways And Transformations Of Dissolved Methane And Dissolved Inorganic Carbon In Arctic Tundra Watersheds: Evidence From Analysis Of Stable Isotopes”. Global Biogeochemical Cycles, 2015, pp. 1893 - 1910.
  • Herndon, E. M., et al. “Pathways Of Anaerobic Organic Matter Decomposition In Tundra Soils From Barrow, Alaska”. Journal Of Geophysical Research: Biogeosciences, 2015, pp. 2345 - 2359.
  • Warren, J. M., et al. “Root Structural And Functional Dynamics In Terrestrial Biosphere Models - Evaluation And Recommendations”. New Phytologist, 2015, pp. 59 - 78.
  • Weston, D. J., et al. “Sphagnum Physiology In The Context Of Changing Climate: Emergent Influences Of Genomics, Modelling And Host-Microbiome Interactions On Understanding Ecosystem Function”. Plant, Cell & Environment, 2015, pp. 1737 - 1751.
  • RoyChowdhury, T., et al. “Stoichiometry And Temperature Sensitivity Of Methanogenesis And Co2 Production From Saturated Polygonal Tundra In Barrow, Alaska”. Global Change Biology, 2015, pp. 722 - 737.
  • Iversen, C. M., et al. “The Unseen Iceberg: Plant Roots In Arctic Tundra”. New Phytologist, 2015, pp. 34 - 58.
  • Devarakonda, R., et al. “Use Of A Metadata Documentation And Search Tool For Large Data Volumes: The Ngee Arctic Example”. 2015 Ieee International Conference On Big Data (Big Data), 2015.


  • Gangodagamage, C., et al. “Extrapolating Active Layer Thickness Measurements Across Arctic Polygonal Terrain Using Lidar And Ndvi Data Sets”. Water Resources Research, 2014, pp. 6339 - 6357.
  • Wullschleger, S. D., et al. “Plant Functional Types In Earth System Models: Past Experiences And Future Directions For Application Of Dynamic Vegetation Models In High-Latitude Ecosystems”. Annals Of Botany, 2014, pp. 1 - 16.
  • Hayes, D. J., et al. “The Impacts Of Recent Permafrost Thaw On Land–Atmosphere Greenhouse Gas Exchange”. Environmental Research Letters, 2014, p. 045005.


  • Hubbard, S. S., et al. “Quantifying And Relating Land-Surface And Subsurface Variability In Permafrost Environments Using Lidar And Surface Geophysical Datasets”. Hydrogeology Journal, 2013, pp. 149 - 169.
  • Wu, Y., et al. “Remote Monitoring Of Freeze–Thaw Transitions In Arctic Soils Using The Complex Resistivity Method”. Vadose Zone Journal, 2013.


  • Lee, H., et al. “Enhancing Terrestrial Ecosystem Sciences By Integrating Empirical Modeling Approaches”. Eos, Transactions, American Geophysical Union, 2012, pp. 237 - 237.
  • McCarthy, H. R., et al. “Integrating Empirical-Modeling Approaches To Improve Understanding Of Terrestrial Ecology Processes”. New Phytologist, 2012, pp. 523 - 525.
  • Graham, D. E., et al. “Microbes In Thawing Permafrost: The Unknown Variable In The Climate Change Equation”. The Isme Journal, 2012, pp. 709 - 712.
  • Xu, C., et al. “Toward A Mechanistic Modeling Of Nitrogen Limitation On Vegetation Dynamics”. Plos One, 2012, p. e37914.


  • Hanson, P. J., et al. “A Method For Experimental Heating Of Intact Soil Profiles For Application To Climate Change Experiments”. Global Change Biology, 2011, pp. 1083 - 1096.
  • Xu, C., et al. “Importance Of Feedback Loops Between Soil Inorganic Nitrogen And Microbial Communities In The Heterotrophic Soil Respiration Response To Global Warming”. Nature Reviews Microbiology, 2011, pp. 222 - 222.
  • Wullschleger, S. D., et al. “Planning The Next Generation Of Arctic Ecosystem Experiments”. Eos, Transactions, American Geophysical Union, 2011, p. 145.


  • Wullschleger, S. D., and M. Strahl. “Climate Change: A Controlled Experiment”. Scientific American, 2010, pp. 78 - 83.